The youngest known tommotiid: Lapworthella bornholmiensis (Poulsen, 1942) from Cambrian Stage 4 to Guzhangian (Miaolingian) strata of Bornholm and southern Sweden

The Camenellan tommotiid Lapworthella bornholmiensis is systematically revised based on the original type material and collections of Small Shelly Fossils at Swedish Museum Natural History. species clearly differentiated from all other by its unique surface ornamentation. Variability in sclerite shape analyzed among 39 specimens four morphotypes are recognized: bilaterally subsymmetrical A sclerites with straight apertural margin subrectangular to subelliptical cross-section; asymmetrical B oblique curved C one concave resulting elliptical sub-triangular cross section; elongated D subcircular section. further divided into two sub-types. allow a detailed comparison lapworthellid taxa we identify pattern recurring across range species, allowing new understanding scleritome structure. stratigraphic includes Cambrian Stage 4 Gislöv Formation, Wuliuan-Drumian Forsemölla Exsulans Limestones Alum Shale Formation Guzhangian limestone fill Proterozoic basement fissures Bohuslän, making this long ranging youngest known fossil record.